系统分类学实验室聚焦纤毛虫原生生物的基础生物学问题，涉及物种多样性、系统与个体发育、区系及分布等。迄今已主持完成了国家自然基金等课题8项和国际科技合作项目2项，在研国家自然基金1项。目前我们特别关注藻食性纤毛虫和专性厌氧纤毛虫的分类与系统学及生态学以及纤毛虫与原核生物的共生作用机制。迄今本组成员及合作者共发表论著200余篇。主要研究方向：

　　纤毛虫的多样性与分类学 主要涉及各类水体及土壤生境中纤毛虫，特别是研究欠缺的盔形纲、斜毛纲、排毛亚纲、前口纲、钩刺纲等类群的分类与物种多样性及区系分布。

　　纤毛虫的细胞发生学 主要揭示纤毛虫在其生活史的二分裂及生理重组期间的皮层纤毛器以及细胞核等结构的发生模式、原基分化及其与老结构的相承关系等。 

　　纤毛虫的系统发育 基于纤毛虫的形态分类及标记基因信息探讨相关类群的系统地位、发生与演化、阶元间的关系。

　　纤毛虫中的共生系统 探讨关键类群纤毛虫与细菌、单胞藻等共生系统的结构、形成与作用机制。

专著/章节

2019 Hu X, Lin X, Song W. Ciliate Atlas: Species Found in the South China Sea. Beijing: Science Press. 631pp（133万字）

2017 李俐琼, 胡晓钟. 条纹小双虫亚型的发生模式。见：宋微波, 邵晨 著 腹毛类纤毛虫的细胞发生模式。北京：科学出版社，2017年12月，498页，第63-69页

2017 樊阳波, 胡晓钟. 成囊双列虫亚型的发生模式。见：宋微波, 邵晨 著 腹毛类纤毛虫的细胞发生模式。北京：科学出版社，2017年12月，498页，第107-112页

2017 胡晓钟, 宋微波. 耳状额斜虫亚型的发生模式。见：宋微波, 邵晨 著 腹毛类纤毛虫的细胞发生模式。北京：科学出版社，2017年12月，498页，第137-142页 

2017 胡晓钟, 宋微波. 柠檬类瘦尾虫亚型的发生模式。见：宋微波, 邵晨 著 腹毛类纤毛虫的细胞发生模式。北京：科学出版社，2017年12月，498页，第167-172页

2017 胡晓钟, 宋微波. 异弗氏全列虫亚型的发生模式。见：宋微波, 邵晨 著 腹毛类纤毛虫的细胞发生模式。北京：科学出版社，2017年12月，498页，第213-218页

2017 胡晓钟. 美丽异角毛虫亚型的发生模式。见：宋微波, 邵晨 著 腹毛类纤毛虫的细胞发生模式。北京：科学出版社，2017年12月，498页，第247-254页

2017 陈旭淼, 胡晓钟. 冠突伪尾柱虫亚型的发生模式。见：宋微波, 邵晨 著 腹毛类纤毛虫的细胞发生模式。北京：科学出版社，2017年12月，498页，第263-269页

2017 芦晓腾, 胡晓钟. 海洋列毛虫亚型的发生模式。见：宋微波, 邵晨 著 腹毛类纤毛虫的细胞发生模式。北京：科学出版社，2017年12月，498页，第305-311页

2017 胡晓钟. 相似拟双棘虫亚型的发生模式。见：宋微波, 邵晨 著 腹毛类纤毛虫的细胞发生模式。北京：科学出版社，2017年12月，498页，第315-320页

2017 胡晓钟, 宋微波. 拟缩颈半腹柱虫亚型的发生模式。见：宋微波, 邵晨 著 腹毛类纤毛虫的细胞发生模式。北京：科学出版社，2017年12月，498页，第379-384页

2017 胡晓钟, 宋微波. 美丽原腹柱虫亚型的发生模式。见：宋微波, 邵晨 著 腹毛类纤毛虫的细胞发生模式。北京：科学出版社，2017年12月，498页，第397-402页 

2009 宋微波, A.沃伦, 胡晓钟. 主编。中国黄渤海的自由生纤毛虫。科学出版社，北京518页。

2003 宋微波, 赵元君, 徐奎栋, 胡晓钟, 龚骏. 海水养殖中的危害性原生动物。科学出版社，北京483页。

1999 宋微波, 徐奎栋, 施心路, 胡晓钟, 类彦立, 魏军, 陈子桂, 史新柏, 王梅. 原生动物学专论。青岛海洋大学出版社，青岛362页。

论文

2024 Feng X., Méndez-Sánchez D., Zhuang W., Li R., Pomahač O., Čepička I., Rotterová J., Hu X. Morphology, morphogenesis, and molecular characterization of Castula specialis sp. nov. (Ciliophora, Armophorea, Metopida). J. Eukaryot. Microbiol. 71: e13014.

2024 Feng X., Zhuang W., Li R, Hu X. Ontogeny and phylogeny of the anaerobic genus Urostomides with supplementing morphological characterization of the type and one little known species (Alveolata, Ciliophora, Armophorea). Zool. J. Linn. Soc. 2024: zlae055.

2024 Jiang L., Sun Q., Zhu X., Hu X. Taxonomy and molecular phylogeny of genus Plagiocampa (Ciliophora, Prostomatea), with redescriptions of two poorly known species. Protist. 175:126059.

2024 Jiang L., Wang C., Liao L., Al-Farraj SA., Warren A., Hu X. Re-evaluation of the systematic position of the order Prostomatida (Protista: Ciliophora), with the establishment of two new genera and two new species. Zool. J. Linn. Soc. 200:849–864.

2024 Kouser F., Liao L., Hu X. Morphology and molecular phylogeny of a Chinese population of Rubrioxytricha guamensis Kumar et al., 2018 (Ciliophora, Hypotrichia). J. Nat. Hist. 58:688–701.

2024 Li R., Zhuang W., Feng X., Zhu X., Hu X. Morphology and molecular phylogeny of three Parasonderia species including a new species (Ciliophora, Plagiopylea). Eur. J. Protistol. 94:126087.

2024 Liao L., Jiang L., Hu X. Integrative data of a novel ciliate (Alveolata, Ciliophora) propose the establishment of Heterodeviata nantongensis nov. sp. BMC Microbiol. 24:27.

2024 Zhuang W., Feng X., Li R., Al-Farraj SA., Hu X. Morphogenesis of an anaerobic ciliate Heterometopus palaeformis (Kahl, 1927) Foissner, 2016 (Ciliophora, Armophorea) with notes on its morphological and molecular characterization. Protist 175:126007.

2023 Ghaffara A., Bai Y., Hu X. First record of two urotsylid ciliates (Protozoa: Ciliophora: Spirotrichea) from Japan expanding the circumscription of species. J. Nat. Hist. 57:724–743.

2023 Jiang L., Wang C., Al-Farraj SA., Hines HN., Hu X. Morphological and molecular examination of the ciliate family Lagynusidae (Protista, Ciliophora, Prostomatea) and descriptions of two new genera and two new species from China. Mar. Life Sci. Technol. 5:178–195.

2023 Li R., Zhuang W., Feng X., Al-Farraj SA., Schrecengost A., Rotterová J., Beinart RA., Hu X. Molecular phylogeny and taxonomy of three anaerobic plagiopyleans (Alveolata: Ciliophora), retrieved from two geographically distant localities in Asia and North America. Zool. J. Linn. Soc. 199:493–510.

2023 Li R., Zhuang W., Feng X., Hines HN., Hu X. First redescription and molecular phylogeny of Trimyema claviforme Kahl, 1933 with the description of a Chinese population of Plagiopyla nasuta Stein, 1860 (Ciliophora, Plagiopylea). Eur. J. Protistol. 90:126003.

2023 Lu B., Hu X., Warren A., Song W., Yan Y. From oral structure to molecular evidence: new insights into the evolutionary phylogeny of the ciliate order Sessilida (Protista, Ciliophora), with the establishment of two new families and new contributions to the poorly studied family Vaginicolidae. Sci. China Life Sci. 66:1535–1553.

2023 Wang C., Jiang L., Liao L., Qu Z., Kumar S., Hu X. Taxonomic and phylogenetic studies on two marine cyrtophorians (Protista: Ciliophora: Phyllopharyngea) with the establishment of a new species. Syst. Biodivers. 21:2269150.

2023 Wang C., Jiang L., Pan H., Warren A., Hu X. New contributions to the Cyrtophoria ciliates (Protista, Ciliophora): Establishment of new taxa and phylogenetic analyses using two ribosomal genes. J. Eukaryot. Microbiol. 70:e12938.

2022 Jiang L., Wang C., Warren A., Ma H., Hu X. New considerations on the systematics of the family Holophryidae (Protozoa, Ciliophora, Prostomatea), with a description of Holophrya paradiscolor sp. nov. and a redescription of Pelagothrix plancticola. Syst. Biodivers. 20:1–15.

2022 Kouser F., Song W., Jiang L., Zhuang W., Wang C., Wu T., Hu X. Taxonomy and phylogeny of two spirotrichean ciliates (Alveolata, Protista, and Ciliophora) from a plateau saline-alkaline lake in Tibet, China, including description of a new species. Front. Mar. Sci. 9:836341.

2022 Li R., Zhuang W., Feng X., Al-Farraj SA., Warren A., Hu X. Phylogeny of the anaerobic ciliate genus Sonderia (Protista: Ciliophora: Plagiopylea), including the description of three novel species and a brief revision of the genus. Mar. Life Sci. Technol. 4:493−512.

2022 Liu M., Liu Y., Zhang T., Lu B., Gao F., Gu J., AlFarraj SA., Hu X., Song W. Integrative studies on the taxonomy and molecular phylogeny of four new Pleuronema species (Protozoa, Ciliophora, Scuticociliatia). Mar. Life Sci. Technol. 4:179–200.

2022 Qu Z., Pan H., Gong J., Wang C., Filker S., Hu X. Historical review of studies on cyrtophorian ciliates (Ciliophora, Cyrtophoria) from China. Microorganisms 10:1325.

2022 Zhuang W., Li R., Feng X., Al-Farraj SA., Hu X. New contribution to the diversity of the anaerobic genus Metopus (Ciliophora, Armophorea), with descriptions of three new marine species. Front. Mar. Sci. 9:884834.

2021 Jiang L., Wang C., Zhuang W., Li S., Hu X. Taxonomy, phylogeny, and geographical distribution of the little-known Helicoprorodon multinucleatum Dragesco, 1960 (Ciliophora, Haptorida) and key to species within the genus. Eur. J. Protistol. 78:125769.

2021 Jiang L., Zhuang W., El-Serehy HA., Al-Farraj SA., Warren A., Hu X. Taxonomy and molecular phylogeny of two new species of prostomatean ciliates with establishment of Foissnerophrys gen. n. (Alveolata, Ciliophora). Front. Microbiol. 12:686929.

2021 Li R., Zhuang W., Wang C., El-Serehy H., Al-Farraj SA., Warren A., Hu X. Redescription and SSU rRNA gene-based phylogeny of an anaerobic ciliate, Plagiopyla ovata Kahl, 1931 (Ciliophora, Plagiopylea). Int. J. Syst. Evol. Microbiol. 71:004936.

2021 Li S., Zhuang W., Pérez-Uz B., Zhang Q., Hu X. Two anaerobic ciliates (Ciliophora, Armophorea) from China: Morphology and SSU rDNA sequence, with report of a new species, Metopus paravestitus nov. spec. J. Eukaryot. Microbiol. 68: e12822.

2021 Liu M., Wang C., Hu X., Qu Z., Jiang L., Al-Farraj Saleh A., El-Serehy Hamed A., Warren A., Song W.B. Notes on Homalogastra similis nom. nov. (Ciliophora, Scuticociliatia), a replacement name for the junior homonym Homalogastra binucleata Liu et al., 2020. Eur. J. Protistol. 81:125841.

2021 Qu Z., Pan H., Wang C., Ma H., Stoeck T., Hu X. Gastronautidae Deroux, 1994 and Trithigmostoma Jankowski, 1967: Evolutionary links among cyrtophorian ciliates (Protista, Ciliophora, Phyllopharyngea). Front. Mar. Sci. 8:625644.

2021 Wang C., Hu Y., Warren A., Hu X. Genetic diversity and phylogeny of the genus Euplotes (Protozoa, Ciliophora) revealed by the mitochondrial CO1 and nuclear ribosomal genes. Microorganisms 9:2204.

2021 Wang C., Jiang L., Qu Z., Al-Farraj SA., Warren A., Hu X. Taxonomy and phylogeny of three species of Dysteria (Ciliophora, Phyllopharyngea) including the description of Dysteria ozakii nom. nov. Protist 172:125831.

2021 Wang R., Bai Y., Hu T., Xu D., Suzuki T., Hu X. Integrative taxonomy and molecular phylogeny of three poorly known tintinnine ciliates, with the establishment of a new genus (Protista; Ciliophora; Oligotrichea). BMC Ecol. Evol. 21:115.

2021 Zhu R., Qu Z., Zhang Q., Filker S., Stoeck T., Li F., Hu X. A new record of Oxytricha granulifera granulifera Foissner and Adam, 1983 (Protozoa, Ciliophora, Oxytrichidae) from a hot spring in Iceland, with notes on its abnormal form during cultivation. Front. Mar. Sci. 8:621349.

2021 Zhuang W., Li S., Bai Y., Zhang T., Al-Rasheid KAS., Hu X. Morphology and molecular phylogeny of the anaerobic freshwater ciliate Urostomides spinosus nov. spec. (Ciliophora, Armophorea, Metopida) from China. Eur. J. Protistol. 81: 125823.

2020 Bai Y., Wang R., Liu W., Warren A., Zhao Y., Hu X. Redescriptions of three tintinnine ciliates (Ciliophora: Tintinnina) from coastal waters in China based on lorica features, cell morphology, and rDNA sequence data. Eur. J. Protistol. 72:125659.

2020 Bai Y. Wang R., Song W., Li L., Santoferrara LF., Hu X. Three redescriptions in Tintinnopsis (Protista: Ciliophora: Tintinnina) from coastal waters of China, with cytology and phylogenetic analyses based on ribosomal RNA genes. BMC Microbiol. 20:374.

2020 Bai Y., Wang R., Al-Rashed KA., Miao M., Hu X. The type species of Amphorellopsis and Tintinnopsis (Protozoa：Ciliophora): a new ciliary pattern and some comments in Tintinnina. J. King Saud Univ. Sci. 32: 3454–3462.

2020 Bai Y., Wang R., Song W., Suzuki T., Hu X. Redescription of five tintinnine ciliates (Alveolata: Ciliophora: Oligotrichea) from coastal waters of Qingdao, China. Mar. Life Sci. Technol. 3:209–221.

2020 Liu M., Wang C., Hu X., Qu Z., Jiang L., Al-Farraj SA., El-Serehy HA., Warren A., Song W. Taxonomy and molecular phylogeny of three species of scuticociliates from China: Citrithrix smalli gen. nov., sp. nov., Homalogastra binucleata sp. nov. and Uronema orientalis Pan et al., 2015 (Protozoa, Ciliophora, Oligohymenophorea), with the proposal of a new family, Citrithrixidae fam. nov. Front. Mar. Sci. 7:604704.

2020 Lu B., Shen Z., Zhang Q., Hu X., Warren A., Song W. Morphology and molecular analyses of four epibiotic peritrichs on crustacean and polychaete hosts, including descriptions of two new species (Ciliophora, Peritrichia). Eur. J. Protistol. 73:125670.

2020 Méndez-Sánchez D., Mayén-Estrada R., Hu X. Euplotes octocarinatus Carter, 1972 (Ciliophora, Spirotrichea, Euplotidae): Considerations on its morphology, phylogeny, and biogeography. Eur. J. Protistol. 74:125667.

2020 Wang R., Song W., Bai Y., Warren A., Li L., Hu X. Morphological redescriptions and neotypification of two poorly known tintinnine ciliates (Alveolata, Ciliophora, Tintinnida), with a phylogenetic investigation based on SSU rRNA gene sequences. Int. J. Syst. Evol. Microbiol. 70:2515–2530.

2019 Li F., Qu Z., Luo D., Filker S., Hu X., Stoeck T. Morphology, morphogenesis and molecular phylogeny of a new obligate halophile ciliate, Schmidtiella ultrahalophila gen. nov., spec. nov. (Ciliophora, Hypotrichia) isolated from a volcanic crater on Sal (Cape Verde Islands). J. Eukaryot. Microbiol. 66:694–706.

2019 Lu B., Li L., Hu X., Ji D., Al-Rasheid K.A., Song W. Novel contributions to the peritrich family Vaginicolidae (Protista, Ciliophora), with morphological and phylogenetic analyses of poorly-known species of Pyxicola, Cothurnia and Vaginicola. Zool. J. Linn. Soc. 187:1–30.

2019 Wang C., Qu Z., Hu X. Morphology and SSU rDNA sequences of four cyrtophorian ciliates from China, with description of a new species (Protista, Ciliophora, Phyllopharyngea). Zootaxa 4664 :206–220.

2019 Wang L., Qu Z., Li S., Hu X. Morphology and molecular phylogeny of two little-known species of Loxodes, L. kahli Dreagesco & Njiné, 1971 and L. rostrum Müller, 1786 (Protist, Ciliophora, Karyotrichea). J. Ocean Univ. China 18:643–653.

2019胡晓钟, 赵研, 黄洁, 高凤, 高珊, 邵晨, 宋微波. 进展中的纤毛虫学研究：新格局、新窗口与新拓展. 海大学报 49:28-42.

2018 Bai Y., Li S., Li Y., Miao M., Hu X. Morphogenesis and molecular characterization of a little known soil ciliate, Oxytricha nauplia Berger et Foissner, 1987 (Ciliophora, Sporadotrichida). Acta Protozool. 57:79–94.

2018 Chen L., Wu W., El-Serehy HA., Hu X., Clamp JC. Morphology, morphogenesis, and phylogeny of an Anteholosticha intermedia (Ciliophora, Urostylida) population from the United States. Eur. J. Protistol. 65:1–15.

2018 Lu B., Ji D., Sheng Y., Song W., Hu X., Chen X., Al-Rasheid KA. New discoveries of the genus Thuricola Kent, 1881 (Ciliophora, Peritrichia, Vaginicolidae), with descriptions of three poorly known species from China. Acta Protozool. 57:123–143.

2018 Méndez-Sáncheza D., Mayén-Estrada R., Luo X., Hu X. A new subspecies of Oxytricha granulifera (Hypotrichia: Oxytrichidae) from Mexico, with notes on its morphogenesis and phylogenetic position. J. Eukaryot. Microbiol. 65:357–371.

2018 Qu Z., Pan H., Lin X., Li L., Aleidan A., Al-Farraj SA., Stoeck T., Hu X. A contribution to the morphology and phylogeny of Chlamydodon, with three new species from China (Ciliophora, Cyrtophoria). J. Eukaryot. Microbiol. 65:236–249.

2018 Song W., Wang L., Li L., Al-Farraj SA., Aleidan A., Smith S., Hu X. Morphological characterizations of four species of Parallelostrombidium (Ciliophora, Oligotrichia), with a note on the phylogeny of the genus. J. Eukaryot. Microbiol. 65:679–693.

2018 Song W., Li J., Huang Y., Hu X., Liu W., Al-Rasheid K., Miao M. Morphology of three aloricate choreotrich ciliates, including description of a new species Parastrombidinopsis costalis sp. n. (Ciliophora, Choreotrichia), and phylogeny of the genus Parastrombidinopsis. Acta Protozool. 57:153–167.

2018 Wang R., Song W., Yi Z., Lin X., Al-Rasheid KA., Hu X. Morphology and molecular phylogeny of two new species of Spirostrombidium (Ciliophora, Oligotrichia), with a key to the species of Spirostrombidium. Syst. Biodivers. 16:743–756.